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The formation of polar cap density enhancements, such as tongues‐of‐ionization (TOIs), are often attributed to enhanced dayside reconnection and convection due to solar wind changes. However, ionospheric poleward moving density enhancements can also form in the absence of changes in the solar wind. This study examines how TOI and patch events that are not triggered by solar wind changes relate to magnetospheric processes, specifically substorms. Based on total electron content and Super Dual Auroral Radar Network (SuperDARN) observations, we find substorms that occur at the same time as TOIs are associated with sudden enhancements in dayside poleward flows during the substorm expansion phase. Active Magnetosphere and Planetary Electrodynamics Response Experiment (AMPERE) observations also show enhanced field‐aligned currents (FACs) that extend into the dayside ionosphere during this period. We suggest that the global enhancement of FACs and convection during these substorms are the drivers of these TOIs by enhancing dayside convection and transporting high‐density lower‐latitude plasma into the polar cap. However, we also find that not all substorms are coincident with polar cap density enhancements. A superposed epoch study showed that the AL index for TOIs during substorms is not particularly stronger than substorms without TOIs, but epoch studies of AMPERE observations do show events with TOIs to have a higher total FAC on both the dayside and nightside. Our results show the importance of TOI formation during substorms when solar wind drivers are absent, and the importance of considering substorms in the global current system. This work also shows the need to incorporate substorms into models of high‐latitude global convection and currents.

Polar cap ionospheric plasma flow studies often focus on large‐scale averaged properties and neglect the mesoscale component. However, recent studies have shown that mesoscale flows are often found to be collocated with airglow patches. These mesoscale flows are typically a few hundred meters per second faster than the large‐scale background and are associated with major auroral intensifications when they reach the poleward boundary of the nightside auroral oval. Patches often also contain ionospheric signatures of enhanced field‐aligned currents and localized electron flux enhancements, indicating that patches are associated with magnetosphere‐ionosphere coupling on open field lines. However, magnetospheric measurements of this coupling are lacking, and it has not been understood what the magnetospheric signatures of patches on open field lines are. The work presented here explores the magnetospheric counterpart of patches and the role these structures have in plasma transport across the open field‐line region in the magnetosphere. Using red‐line emission measurements from the Resolute Bay Optical Mesosphere Thermosphere Imager, and magnetospheric measurements made by the Cluster spacecraft, conjugate events from 2005 to 2009 show that lobe measurements on field lines connected to patches display (1) electric field enhancements, (2) Region 1 sense field‐aligned currents, (3) field‐aligned enhancements in soft electron flux, (4) downward Poynting fluxes, and (5) in some cases enhancements in ion flux, including ion outflows. These observations indicate that patches highlight a localized fast flow channel system that is driven by the magnetosphere and propagates from the dayside to the nightside, most likely being initiated by enhanced localized dayside reconnection.